FRANCE: Berlese (1916b); Ryke (1962a);

HUNGARY: Kandil (1983);

LATVIA: Lapinya (1976b); Salmane (1999); Salmane (2000); Salmane (2003);

 

Substrate: From moss — Berlese (1916b), Ryke (1962a); From nest of bird — Lapinya (1976b); From soil — Kandil (1983), Salmane (2000), Salmane (1999), Salmane (2003);